The following is a detailed comparison between kindle and the Sony Reader(PRS-505).

Sony E-Reader Kindle

Summary

Screen: Sony is better – 8 level gray scale as opposed to Kindle’s 6.

Size: Sony is smaller in every dimension.

Computer Requirement: Required at all times for sony content download. Need sometimes for Kindle (only in non-wireless coverage areas).

Format:
Sony – AA, PRC, MOBI, AZW, HTML not covered
Kindle – BBeB, AAC, RTF not covered
everything else equal

Expansion: Sony offers more capacity to expand and hold additional content – up to 10 GB.

Connectivity: Both utilizes computer as a connectivity source. Kindle bypasses this requirement when in a wireless coverage area.

Content: Much more content availability with Sony.

Weight: Sony reader a little lighter.

Capacity: Kindle holds more titles in internal memory.

Battery Power: Both devices give good content mileage.

Fees: Neither device charges a fee for download.

Wikipedia Access: Both can deliver content. Kindle can do wirelessly.
Accessories: Just about the same (Sony includes connection software).

Conclusion:
Sony has a better screen, is smaller, requires computer as mediator for content download, delivers more available content, weighs less, charges no fees, and can deliver wikipedia content.

Kindles holds more in internal memory, delivers its contents wireless when in coverage areas otherwise computer mediator needed.

So which device do you choose? Your choice really depends on your taste.

If you are looking for a device that delivers content wirelessly and hold more in internal memory, is a reasonable size and weight, and give access to a decent amount of diverse content, then you may want to choose Kindle.

If you are looking for a device that does not necessarily deliver content wirelessly, but still delivers it in a timely manner, holds a decent amount of content in internal memory, gives access to a huge content inventory, is a reasonable size and weight (smaller and lighter than Kindle), then Sony may be your choice.

Regardless of which device you choose, you will be getting one of the best electronic book readers on the market. Good Luck on your choice

Update I
Can you Rotate the screen?
YES for the PRS-505, NO for the kindle . To rotate the Screen on your PRS-505, simple hold down the “Size” button for about 5 seconds and the screen will rotates from portrait to landscape and back.

Can you jump forward more than one page at a time in either device?
YES for both devices.
For the PRS-505, hold the “Page Forward” or “Back” button for five seconds and the page will jump 10 pages in the direction of the button. For the Kindle, hold down the “Alt + Prev Page” or “Alt + Next Page” buttons and the page will jump in the direction of the button by quite a few pages. I am not sure how much, but it seems to be a little more than the PRS-505

* For the PRS-500:”1″ brings you to the start of the book. “2″ brings you to 20% of the book. “3″ to 30% and so on. So if you want to open to about half way in the book simply hit “5″.

How Can I Make PDF Look Better on the Device?
For the Kindle , you cannot do much right now – PDF display is still in the experimental phase. As for the PRS-505, its just a simple matter of reformatting your PDF document to fit the device. Most PDF documents are formatted to fit an 8 ½” x 11″ piece of paper and not a screen the size of the Reader. In this case the Reader will have to resize your files to display on its screen. Your files may look better if you reformat them with the Reader’s screen size in mind. You can go here to download directions on how to optimize PDFs for the Reader.

Why does the page sometimes change when I place the device in my bag?
You can control this for both Kindle and the PRS-505:
For the Kindle, simple press “Alt+A” to put the device to sleep before you put it away.
For the PRS-505, you will need to turn the device off using the power switch on the left hand side of the reader which will in-turn lock the buttons. When you are ready to read again, just turn the unit on and you will pick up right where left off.

Can You Lock the Device?
For the PRS -505, you can lock the device by putting on a four digit password that will need to be entered anytime the Reader is turned on. If you use this feature you will have to remember your password or call 1-866-962-7669.
No such password access for Kindle.

Update II – Listening to Audio files

Kindle
You can download (via USB cable) and listen to MP3 files only as background music. This means that you cannot create a play list or organize the files the way you would on a MP3 player. Once the music starts it will play until you stop it or switch to audio books. You can adjust the volume using the built-in volume control at the bottom of the device.

PRS-505
You can download and listen to MP3 files on the Sony reader as you would on the player itself or as background music while reading. Tracks are listed in alphabetical order. You simply need to select the music of choice, press “ENT,” and you are on your way. In the play mode, you can see “everything:” track name, artist name, album name, elapsed time, track number, total number of track, etc. You can fast forward, rewind, stop, play, adjust volume the normal way with the built-in volume control on the device.

Speakers
You have the option of listening to audio file on kindle via the external built-in speakers or privately using a headphone. PRS-505 gives you no choice but to listen with a headphone as It is not equipped with external speakers.

Both devices use up a significant amount of battery power when audio is played…so use conservatively.

Audiobooks
If you prefer to listen to your books as opposed to reading them, Kindle gives you the option of downloading and listening to audio books from Audible.com. PRS-505 does not offer you this option. For more information on how to access audio books for the Kindle go here.

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You Can Leave Your Thoughts

Do you like to take notes and make comments while reading? You can do this with your kindle. All you have to do is highlight lines of text to type in notes to any of the content on Kindle. Amazon will automatically stores all of your annotations in the “My Clippings” file and even backs them up on Amazon servers so they will never be lost even if you lose or damage your Kindle.

Once you highlight your text, a small note icon will appear to the right of the text. Use the select wheel to move the cursor to the note icon then press the select wheel and choose “Edit Note” from the menu. Now move the cursor to the line where you want to add your thoughts, press the select wheel, and choose “Add Note” from the menu. Use the keyboard to type your note then select “Done”. Later you can view the note, edit it, remove it, or see all of your annotations in something you are reading by selecting “My Notes & Marks” from the menu.